The story picks up where Nikolai and his son, Innokenti live in a secluded house in America. One day they’re approach by a man named Sergei who states that the mentor has demanded Niklaoi’s return to Russia.

On the other hand, Daniel Cross in present day is taken off the Animus for his own well-being, yet he still experiences Nikolai’s memories through the Bleeding Effect. We follow his journey as he gets inducted to the Inner Sanctum of the Templar Order and makes a shocking discovery.